2025-2026 Graduate Catalog
Biology
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Mark Ungerer, Director
Michael Veeman, Director of graduate studies
116 Ackert Hall
785-532-6615
Fax: 785-532-6653
mcungere@ksu.edu
http://www.ksu.edu/biology/
For more detailed information on faculty research interests, see http://www.k-state.edu/biology/people/tenure/index.html.
Programs
Biology and Microbiology are broad and innovative disciplines, and our graduate program reflects both of these characteristics. Our faculty members are research scientists with a breadth and depth of research expertise, and they apply that expertise in their roles as graduate mentors. The graduate faculty members of the Division of Biology strongly believe that independent research is the preeminent feature of our graduate programs. Research expertise represented within the Division of Biology ranges from studies of cell development and cell signaling to the influence of grazers and fire with plant and animal communities on the tallgrass prairie. Graduate opportunities are available in all areas of biology, including molecular, cellular and developmental biology; community, ecosystem and landscape ecology; wildlife biology; cancer biology; microbiology; virology; immunology; parasitology; and genetics.
In partnership with a major professor and a supervisory committee, each graduate student develops an individualized program of study, consisting of course work and laboratory or field research. Each program of study is tailored to meet the specific needs of the individual student. Unlike more formal curricula with rigid requirements, our flexible approach allows students to pursue their professional goals more efficiently and more effectively. All beginning graduate students are required to take BIOL 863 (Professional Skills in Biology), where they learn about grant-writing, presentation skills, research ethics, effective instructional techniques, and other skills that will be useful to them throughout their careers as professional scientists. This course is the only required course; all other courses are chosen in consultation with the advisory committee. A minimum of 30 credit hours beyond the bachelor’s degree is required to earn a master of science degree; a minimum of 90 credit hours beyond the bachelor’s degree is required to earn a doctoral degree. In addition, we require that each student write a thesis or dissertation based on their original research; this work should be of sufficient quality and importance to merit publication in a refereed journal. For additional details regarding responsibilities and expectations for our graduate students, see http://www.ksu.edu/biology/grad/responsibilities.html.
Facilities
Graduate research and education takes place primarily in four on-campus sites–Ackert Hall, Bushnell Hall, Chalmers Hall, and Leasure Hall–as well as at the off-campus site of the Konza Prairie Biological Station.
Ackert Hall
Ackert Hall is the main biology facility, containing 130,000 square feet of research and instructional space. Ackert Hall provides modern, well-equipped laboratories for teaching and research, and, contains the Division offices, a rooftop greenhouse, an electronics shop, a research supply storeroom, and animal care facilities. Equipment available for research includes ultracentrifuges, growth chambers, a research microscope facility (with confocal and transmission electron microscope plus image processing capabilities), mass spectroscopy facilities, and many other research and support equipment items that are required for a modern biology research program. Research labs overseen by faculty members in all three of our sections (see above) can be found in Ackert Hall.
Bushnell Hall
Graduate faculty members with an interest in ecological studies have offices and laboratory space in Bushnell Hall. Included in this building are the herbarium, environmental chambers, dark rooms, analytical laboratories, and tanks for fish culture ranging from 100- to 2,000-liter capacity. Extensive computing and imaging facilities support the climatologic and remote sensing activities.
Chalmers Hall
Chalmers Hall was completed in November 2002 and is adjacent to Ackert Hall. This modern facility houses ten Division of Biology faculty and research programs in 12,300 square feet on the second floor of the building. Faculty members with laboratories in Chalmers Hall have research interests in developmental biology, immunology, virology, bioinformatics, ecological genomics, and vector biology.
Leasure Hall
Leasure Hall houses the Kansas Cooperative Fish and Wildlife Research Unit. This unit is a partnership of the U.S. Department of the Interior, Kansas State University, the Kansas Department of Wildlife and Parks, and the Wildlife Management Institute. Two U.S. Department of the Interior employees, with appointments as Division of Biology adjunct graduate faculty members, oversee fisheries and wildlife research programs as part of their duties in the Kansas Cooperative Fish and Wildlife Research Unit.
Konza Prairie Biological Station
The Konza Prairie Biological Station is located only a few miles south of the main KSU campus, allowing for ease of access for faculty members and graduate students working there. This station contains approximately 8,616 acres of native tall grass prairie, and has been set aside by the Nature Conservancy as a unique outdoor laboratory for long-term research. This land was purchased with funds provided by Katharine Ordway, and leased to the Division of Biology by the Nature Conservancy. Experimental manipulations of the landscape, involving grazing and/or burning, allow researchers to focus on patterns and processes which might be important in maintenance of the tall grass prairie ecosystem.
Research strengths
The graduate faculty members of the Division of Biology have a proven track record in scientific achievement, and they continue to excel in competition for extramural research funding. Our research programs are funded by the state of Kansas, via support from the Kansas Agricultural Experiment Station and Kansas Wildlife and Parks, and by direct grants to our faculty members from a diverse group of agencies and foundations, including the National Science Foundation, the National Institutes of Health, the Environmental Protection Agency, the Department of Interior, the Department of Energy, DOD, the American Heart Association, the American Cancer Society, as well as others. Areas of particular research strength, which provide collaborative research opportunities for our graduate students, include:
Long-Term Ecological Research
The National Science Foundation has supported Division of Biology research activities on the Konza Prairie continuously since 1980. This long-term approach is designed to investigate gradual and subtle changes in microbe, plant and animal diversity and numbers on the tall grass prairie, the impact of human activities on ecosystem productivity, and changes in ecological processes caused by burning and/or grazing.
Kansas Lipidomics Research Center
The Kansas Lipidomics Research Center is a collaborative venture of scientists in the Division of Biology, in the Department of Biochemistry and Molecular Biophysics, and at the University of Kansas. The Center performs comprehensive, quantitative profiling of lipid molecular species with high sample throughput, using mass spectrometry and other innovative technologies. Diversity and concentrations of lipid metabolites in genomically altered and physiologically manipulated organisms are investigated in order to identify the functions of genes that encode proteins involved in lipid metabolism. The Center is also dedicated to improving lipid analytical technologies, to promoting collaborative research among lipid scientists, and to providing training opportunities for post doctoral, graduate, and undergraduate students. More information is available at http://www.k-state.edu/lipid/lipidomics/.
Ecological Genomics
The Ecological Genomics initiative is an interdisciplinary research enterprise, aimed at exploring the genetic bases for organismal responses to the environment. Students involved in the program engage in a truly multidisciplinary program, working with faculty members in ecology and molecular biology/genetics. These research efforts employ functional genomic techniques to probe the interface between ecology and genetics. More information about the Kansas Ecological Genomics collaborative research groups can be found at http://www.ksu.edu/ecogen/.
Other multi-investigator research initiatives
Information about many other exciting collaborative research efforts can be found at https://www.k-state.edu/biology/research/interdisciplinary.html
Admission and financial support
For detailed information on admission and financial support, see http://www.k-state.edu/biology/grad/index.html.
Accelerated
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Accelerated Biology - Animal Biology (BS)/Biology (MS)
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Accelerated Biology - Cellular and Molecular Biology (BS)/Biology (MS)
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Accelerated Biology - Computational Biology (BS)/Biology (MS)
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Accelerated Biology - Ecology and Evolutionary Biology (BS)/Biology (MS)
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Accelerated Biology - Human Health Biology (BS)/Biology (MS)
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Accelerated Biology - Integrative Biology (BS)/Biology (MS)
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Accelerated Biology - Plant Biology (BS)/Biology (MS)
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Accelerated Fisheries, Wildlife, Conservation, and Environmental Biology – Conservation Biology(BS)/Biology(MS)
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Accelerated Fisheries, Wildlife, Conservation, and Environmental Biology – Environmental Biology(BS)/Biology(MS)
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Accelerated Fisheries, Wildlife, Conservation, and Environmental Biology – Fisheries Ecology and Management(BS)/Biology(MS)
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Accelerated Fisheries, Wildlife, Conservation, and Environmental Biology – Wildlife Ecology and Management(BS)/Biology(MS)
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Accelerated Microbiology – General Microbiology (BS)/Biology(MS)
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Accelerated Microbiology – Medical Microbiology(BS)/Biology(MS)
Doctor of Philosophy
Master of Science
Biology
-
BIOL 100 - Biology First-Year Orientation Credits 1
Each meeting of Biology Freshman Seminar covers a topic related to student success, including student help sources such as tutoring and academic coaching, the Career Center, student involvement opportunities within the Division of Biology (such as research), the university, and community. In addition, students develop a graduation plan after going over degree requirements and gain competency in navigating KSU online resources.
Repeat for Credit
N
Typically Offered
Fall
-
 BIOL 101 - Concepts of Biology Credits 4
Fundamental concepts in evolutionary biology, ecology, cell biology, genetics, biodiversity, and the scientific method. The role of biology in society and the development of skills aimed at differentiation of biological fact from fiction are emphasized and explored.
Note
Concepts of Biology is an introductory biology course with a lab, and is intended for non-science majors.
Repeat for Credit
N
Typically Offered
Fall, Spring
K-State 8
Natural and Physical Sciences
-
BIOL 198 - Principles of Biology Credits 4
An introductory biology course with emphasis on understanding processes and linkages within and between different levels in the hierarchy of life. Specific areas covered include the scientific method, evolution, ecology, biological molecules, structure/function of cells, genetics, energy flow, biodiversity, and plant/animal biology. BIOL 198 is required in many science degree programs.
Note
Two two-hour studio sessions incorporating lecture and lab elements. At least one year of high school biology or chemistry is recommended.
Repeat for Credit
N
Typically Offered
Fall, Spring, Summer
K-State 8
Empirical and Quantitative Reasoning
K-State 8 Tag 2
Natural and Physical Sciences
-
BIOL 221 - Organismic Biology Credits 5
A study of the structure and function of organisms with special attention paid to the phylogenetic origins of taxonomic groups and the integration of their structural systems.
Note
Three hours lecture and four hours lab.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 198 or equivalent.
Typically Offered
Fall, Spring
K-State 8
Natural and Physical Sciences
-
BIOL 245 - Principles of Histology Credits 2
Introduction to the field of histology including methods of sample preparation, embryonic and fetal development of tissues, structure of primary tissues in fully formed vertebrate body, and examination of those primary tissue types in a variety of organs.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 198 or equivalent recommended.
Typically Offered
Summer
-
BIOL 255 - General Microbiology Credits 4
Microorganisms; their handling, morphology, growth, and importance.
Note
Two hours lecture and four hours lab a week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 198 and one course in chemistry.
Typically Offered
Fall, Spring
K-State 8
Empirical and Quantitative Reasoning
K-State 8 Tag 2
Natural and Physical Sciences
-
BIOL 263 - Ecology of Environmental Problems Credits 3
Principles of ecology and their application to such problems as pollution, human population growth, and land-use planning.
Note
Two hours lecture and one hour discussion a week.
Repeat for Credit
N
Requisites
Prerequisite: One course in natural science.
Typically Offered
Fall
K-State 8
Global Issues and Perspectives
K-State 8 Tag 2
Natural and Physical Sciences
-
BIOL 330 - Public Health Biology Credits 3
Fundamental concepts of human infectious and organic diseases with emphasis on disease etiology and mechanisms, collection of epidemiological data, and the influences upon, and consequences of, governmental public health policy.
Note
Two hours lecture and one hour recitation per week.
Repeat for Credit
N
Requisites
Prerequisite: One course in a natural science.
Typically Offered
Fall
K-State 8
Global Issues and Perspectives
K-State 8 Tag 2
Natural and Physical Sciences
-
BIOL 333 - Introduction to Fisheries, Wildlife, Conservation and Environmental Biology Credits 3
An introduction to the ecological and management principles associated with the fields of fisheries, wildlife, conservation, and environmental biology. Topics include population ecology and assessment, resource management, and environmental legislation. Prepares FWCEB majors for more advanced coursework and promotes the understanding of natural resource management for non-majors.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 198.
Typically Offered
Spring
-
BIOL 365 - Practicum in Biology Credits 1-4
Supervised experience as a teaching assistant in biology courses. Approximately 3 hours of commitment per week is expected for each credit hour earned. One hour recitation a week plus three to nine hours lab a week.
Note
Permission of instructor and credit with superior performance in the course in which the student will be involved.
Repeat for Credit
Y
Typically Offered
Fall, Spring, Summer
-
BIOL 397 - Topics in Biology Credits 1-6
Special course offering in an area of faculty expertise and/or supervised independent study project.
Repeat for Credit
Y
Typically Offered
Fall, Spring, Summer
-
BIOL 398 - Principles of Genetics Credits 4
An introduction to classical, molecular, quantitative, and population genetics. Human examples and comparisons between prokaryotic and eukaryotic genetics are emphasized. Topics include Mendelian inheritance, DNA and chromosome structure, cell division and DNA replication, gene expression, mutation, recombinant DNA, genomics, quantitative inheritance, and population/evolutionary genetic principles.
Note
Three hours lecture and one hour recitation.
Repeat for Credit
N
Requisites
Prerequisites: BIOL 198 and CHM 210 OR CHM 220 or concurrent enrollment. Concurrent enrollment in or completion of a college-level math or statistics course is recommended.
Typically Offered
Fall, Spring
K-State 8
Empirical and Quantitative Reasoning
K-State 8 Tag 2
Natural and Physical Sciences
-
BIOL 399 - Honors Seminar in Biology Credits 1-3
Selected topics.
Note
Open to non-majors in the honors program.
Repeat for Credit
Y
Typically Offered
Spring
-
BIOL 410 - Biology of the Cancer Cell Credits 3
Current concepts of cancer biology including roles of mutation, cell division, apoptosis, stem cells, microbes, and the immune system in promoting and/or inhibiting cancer.
Repeat for Credit
N
Requisites
Prerequisite: Two courses in biology (BIOL 398 recommended).
Typically Offered
Fall
K-State 8
Natural and Physical Sciences
-
BIOL 432 - Wild Bird Research Credits 3
Hands-on training in both field methods involving the capture, handling, measurement, and sampling of wild birds as well as training in data manipulation, analysis in R, visualization, collaboration, and presentation of original research. Requires weekend participation once monthly. Assessment based on completion of assignments and project components as well as effort and progress in learning field methods.
Note
Note – 2-hour, Friday morning class sessions weekly; 3 consecutive mornings (Fri-Sat-Sun) in the field each month. Consent of instructor
Repeat for Credit
N
Typically Offered
Spring
-
BIOL 435 - Environmental Biology Credits 3
An exploration of biological aspects, particularly ecological and evolutionary, as related to environmental issues. Topics include toxicology, global environmental issues, eutrophication, genetically modified organisms, emerging diseases, conservation, and relationship of human nature to causes of and solutions for environmental issues.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 198, CHM 110 or 210.
Typically Offered
Spring
-
BIOL 441 - Human Body I Credits 4
Human anatomy with an introduction to human physiology. Examination of cadavers, dissection of specimens, use and interpretation of EKGs, myelograms, and radiographs.
Note
Three hours of lecture and two hours of lab per week.
Repeat for Credit
N
Requisites
Prerequisites: BIOL 198, CHM 110/CHM 111 OR CHM 210, BIOL 398 Principles of Genetics OR PHYS 214 Engineering Physics II OR KIN 220 Biobehavioral Basis of Physical Activity OR any biology course with a lab component and BIOL 198 as a prerequisite, AND Sophomore standing. Recommended: Strongly recommended that students receive Bs or better in all prerequisite courses. It is also strongly recommended that transfer students complete at least one K-State Biology class before enrolling in BIOL 441.
Typically Offered
Fall, Spring
-
BIOL 442 - Human Body II Credits 4
Human physiology with a review of anatomy from Human Body I.
Note
Three hours of lecture and two hours of lab per week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 441.
Typically Offered
Fall, Spring
-
BIOL 443 - Applied Anatomy of Cadavers Credits 3
Course with lab components and teaching support in Human Body I (HB I; BIOL 441) lab. Consists of brief lectures, dissections, reviewing material, exams, a final (autopsy report), and teaching support for anatomical demonstrations to HB I students. Objectives are: to develop cadaver dissection competence, to aid in a deeper understanding of anatomy and physiology, and to enhance critical thinking and teaching efficacy skills.
Repeat for Credit
N
Requisites
Prerequisite: Have earned a B or better in Human Body I (BIOL 441).
Typically Offered
Fall, Spring
-
BIOL 461 - Phage Hunters I Credits 3
Isolate bacteriophages from soil samples collected. Conduct electron microscopy, isolate bacteriophage DNA, and submit bacteriophage genomes for sequencing.
Note
One hour lecture and three hours lab per week.
Repeat for Credit
N
Typically Offered
Fall
-
BIOL 462 - Phage Hunters 2 Credits 2
Analyze the genomes through bioinformatics and submit the collected data to Howard Hughes Medical Institute (HHMI) and GenBank.
Note
One hour lecture and three hours lab per week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 461
Typically Offered
Spring
-
BIOL 495 - Topics in Biology Credits 1-6
Supervised independent study project and/or special course offering in an area of faculty expertise.
Note
Consent of instructor
Repeat for Credit
Y
Typically Offered
Fall, Spring
-
BIOL 496 - Honors Tutorial in Biology Credits 1-3
Individual directed research and study of a topic in biology, normally as a prerequisite to writing a senior honor thesis.
Repeat for Credit
N
Requisites
Prerequisite: Sophomore standing, membership in the honors program of the College of Arts and Sciences, and permission of instructor.
Typically Offered
Spring
-
BIOL 497 - Honors Project Credits 3
Open only to Arts & Sciences students who are active members of the University Honors Program.
Repeat for Credit
Y
Typically Offered
Fall, Spring, Summer
-
BIOL 500 - Plant Physiology Credits 3
Detailed consideration of physiological processes of higher plants.
Note
Three hours lecture a week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 198 and BIOCH 265 or another course in organic chemistry.
Typically Offered
Fall
-
BIOL 501 - Plant Physiology Lab Credits 1
Lab related to physiological processes of higher plants.
Note
Three hours of lab a week.
Repeat for Credit
N
Requisites
Prerequisite: Concurrent or previous enrollment in BIOL 500.
Typically Offered
Fall
-
BIOL 504 - Plant Ecology Credits 3
Study of the diversity and distribution of vegetation, relationships of plants to their physical environment, plant-animal-microbe interactions, population structure and dynamics, and plant community structure diversity, and succession. Field exercises focusing on methods of grassland and forest vegetation sampling and analysis.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 198 and BIOL 221 or equivalent.
Typically Offered
Fall
K-State 8
Natural and Physical Sciences
-
BIOL 510 - Developmental Biology Credits 3
Introduction to the stages and mechanisms of embryonic animal development. Integrated approach that includes classic experimental embryology and the genetic and molecular regulation of invertebrate and vertebrate animal development.
Note
Three hours lecture per week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 398.
Typically Offered
Spring
-
BIOL 511 - Developmental Biology Laboratory Credits 1
Developmental Biology Laboratory
Note
One three-hour lab a week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 510 or concurrent enrollment.
Typically Offered
On sufficient demand
-
BIOL 513 - Physiological Adaptations of Animals Credits 4
Integration of physiological mechanisms as the basis for adaptive responses of animals to different environments.
Note
Three hours lecture and two hours lab a week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 221; and a course in organic chemistry or biochemistry.
Typically Offered
Fall
-
BIOL 515 - Behavioral Ecology Credits 3
Study of the social, environmental, genetic, and evolutionary processes that affect animal behavior. Topics include: evolution of social organization, spacing and group behavior, mating systems and parental care, sexual selection, communication, aggression, habitat selection, and foraging. Research project required.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 198.
Typically Offered
Spring
-
BIOL 520 - Evolution Credits 3
A study of the theory of evolution including its historical and social implications.
Note
Three hours lecture a week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 398 or a course in genetics.
Typically Offered
Spring
-
BIOL 521 - Evolution of Infectious Diseases Credits 3
The interplay of ecological interactions, evolutionary dynamics, and the mechanisms of host-microbe and microbe-microbe interactions shape the emergence and impact of infectious diseases. An interactive approach examines relationships among these factors. Key topics of discussion will include zoonotic infections, emerging infectious diseases, the spread of antimicrobial resistance, and the impact of public health measures.
Note
Three hours lecture a week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 398 or BIOL 255 or ASI 500.
Typically Offered
Spring
-
BIOL 529 - Ecology Credits 3
Interdisciplinary examination of organisms and their interaction with the environment, ecosystem structure and function, population ecology and demography, community structure and dynamics, and basic ecological principles and their relevance to contemporary environmental issues.
Note
Three hours lecture per week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 198 and CHM 210.
Typically Offered
Fall, Spring
-
BIOL 530 - Pathogenic Microbiology Credits 3
Etiology and descriptions of major infectious diseases of humans within the perspective of host defenses.
Note
Two hours lecture and one hour laboratory-demonstration a week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 255.
Typically Offered
Fall
-
BIOL 541 - Cell Biology Credits 3
Structure and function of cells and subcellular components. A molecular understanding of membranes and cellular physiology will be emphasized.
Note
Three hours lecture.
Repeat for Credit
N
Requisites
Prerequisite: (BIOL 398 or ASI 500) and (CHM 350 or CHM 531).
Typically Offered
Fall, Spring
-
BIOL 542 - Ichthyology Credits 3
Systematics, morphology, physiology, distribution, and natural history of fishes.
Note
Two hours lecture and three hours lab a week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 221.
Typically Offered
Spring-even years
-
BIOL 543 - Ornithology Credits 3
Classification, morphology, physiology, distribution, and natural history of birds.
Note
Two hours lecture and three hours lab a week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 221
Typically Offered
Spring
-
BIOL 544 - Mammalogy Credits 3
Characteristics, evolution, life histories, and ecology of mammals, especially North American game species.
Note
Two hours lecture and three hours lab a week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 221.
Typically Offered
Fall
-
BIOL 545 - Human Parasitology Credits 3
Protozoan and helminth parasites of humans with lesser emphasis on ectoparasitic arthropods. Emphasis on life cycles, control, and laboratory diagnosis.
Note
Three hours lecture a week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 198.
Typically Offered
Spring
-
BIOL 546 - Human Parasitology Laboratory Credits 1
Examination of prepared materials and identification of internal parasites of man.
Note
Two hours lab a week.
Repeat for Credit
N
Requisites
Prerequisite: Concurrent enrollment in BIOL 545.
Typically Offered
Spring
-
BIOL 551 - Taxonomy of Flowering Plants Credits 4
Morphology, taxonomy, and biogeography of the vascular plants.
Note
Two hours lecture and two three-hour labs a week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 221.
Typically Offered
Fall
-
BIOL 580 - Molecular Biology of Genes and Genomes Credits 3
The molecular biology of eukaryotic gene expression and genome function, with an emphasis on gene expression as a regulated process. Selected topics in recombinant DNA and biotechnology, focusing on topics central to a modern understanding of gene regulation.
Note
Three hours lecture a week.
Repeat for Credit
N
Requisites
Prerequisites: BIOL 398 and CHM 350.
Typically Offered
Fall
-
BIOL 604 - Biology of the Fungi Credits 3
An introduction to fungal structure, function, physiology, ecology, and genetics. Importance of fungi as disease organisms, as saprotrophs, and in industry. Techniques of isolation, cultivation, and as experimental organisms.
Note
Two hours lecture and two hours lab a week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 198.
Typically Offered
Fall
-
BIOL 607 - Human Genetics Credits 3
An exploration of human genetics from a molecular perspective, grounded in clinical applications. Provides theoretical knowledge and practical skills in human genome analysis, emphasizing the analysis of case studies and molecular diagnostics techniques. Through a combination of lectures, hands-on laboratory exercises, and discussions, students will examine the implications of genetics and genomics for medicine, clinical diagnostics, genetic counseling, and related fields.
Note
One hour of lecture and two two-hour lecture/labs per week.
Repeat for Credit
N
Requisites
Prerequisites: BIOL 398 Principles of Genetics OR ASI 500 Genetics, and Junior standing.
Typically Offered
Spring
-
BIOL 609 - Cellular and Molecular Biology of Human Diseases Credits 3
Fundamental basis of the major common non-infectious diseases and disorders affecting our society, with emphasis on the biochemical and molecular biological mechanisms by which the structures and functions of specific human tissues, organs and systems are altered.
Note
Three hours lecture per week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 398 and BIOCH 521.
Typically Offered
Fall
-
BIOL 612 - Freshwater Ecology Credits 4
Basic ecological principles of aquatic environments and environmental applications. Plants and animals of local streams, rivers, ponds, and reservoirs are used to demonstrate the interaction of biological processes with the chemical and physical features of natural aquatic environments.
Note
Three hours lecture, three hours lab a week; two optional weekend field trips.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 198 and CHM 110 or CHM 210.
Typically Offered
Fall-Even Years
-
BIOL 632 - Ecology Laboratory Credits 1
Laboratory and field experiences with ecological problems.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 529 and STAT 240 or equivalent.
Typically Offered
Spring
-
BIOL 636 - Cancer Therapy Credits 3
Current methods of cancer management with emphasis on the kinetic principles of chemotherapy and radiation therapy; diagnosis; surgical oncology; oncologic emergencies; adverse effects of cancer therapy; and the new therapies.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 398 and BIOCH 521.
Typically Offered
Spring
-
BIOL 640 - Population Biology Credits 3
An introduction to the theories of quantitative population biology including dynamics, demography and genetics. Emphasis on spatio-temporal variation within and among populations and species.
Note
Three hours lecture a week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 398 and BIOL 529 and STAT 240.
Typically Offered
Fall
-
BIOL 642 - Principles of Conservation Biology Credits 3
Biological diversity and the factors contributing to loss of biodiversity. Scientific principles of biological conservation emphasizing the application of ecological theory and population genetics to the conservation of threatened populations, species and ecosystems.
Note
Three hours lecture per week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 398 and BIOL 529.
Typically Offered
Spring
-
BIOL 670 - Immunology Credits 3
Covers history and overview of the mammalian immune system, innate immunity, immunoglobulins, cytokines, immunological methods, complement system, T cell and B cell development and their differentiation, barrier immunity, vaccines, hypersensitivity reactions, autoimmunity, immunodeficiencies, and anticancer immunity.
Repeat for Credit
N
Requisites
Prerequisite: BIOCH 521 and BIOL 398 or ASI 500 with a C or better. BIOL 255 and BIOL 541 and senior standing are highly recommended.
Typically Offered
Spring
-
BIOL 671 - Immunology Lab Credits 2
Laboratory exercises in immunology.
Note
Three-hour lab a week plus one hour recitation.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 670 or concurrent enrollment.
Typically Offered
Spring
-
BIOL 675 - Genetics of Microorganisms Credits 3
The genetics of bacteria, viruses, and other microorganisms. Both the use of genetics in microbiological studies and the use of microbial systems to investigate basic genetic problems will be covered.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 398 and BIOL 255.
Typically Offered
Fall
-
BIOL 676 - Molecular Genetics Laboratory Credits 3
An advanced course in the techniques of molecular genetics and recombinant DNA technology, including molecular cloning, library screening, polymerase chain reaction, and RFLP and DNA sequence analysis. A number of projects will be compelted during the semester.
Note
One-hour lecture and two three-hour labs.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 675 or concurrent enrollment.
Typically Offered
Fall
-
BIOL 682 - Fish Ecology Credits 3
The interaction between fish and their environment. Exploring fundamental ecological processes in aquatic systems at individual, population, community and ecosystem scales.
Note
Two hours lecture and three hours lab per week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 529.
Typically Offered
Spring-Even Years
-
BIOL 684 - Wildlife Management and Techniques Credits 4
Concepts and techniques of wildlife ecology and management with emphasis on North American game species. Applied population dynamics as they relate to historical and recent developments in wildlife management, habitat improvement, and related material.
Note
Three hours lecture, one hour recitation, and three hours lab a week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 333.
Typically Offered
Spring
-
BIOL 687 - Microbial Ecology Credits 3
The ecology of aquatic, terrestrial, animal and plant host-associated microorganisms in their natural environments.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 255 or BIOL 529.
Typically Offered
Spring
-
BIOL 695 - Internship in Biology Credits 1-3
Undergraduate internship in Biology pursued outside Kansas State University, allowing the student to gain experience in career-relevant work. A minimum of 45 hours spent in internship is expected for each credit hour in which a student is enrolled
Note
This offering is coordinated through Biology’s Office of Undergraduate Studies, and arrangements for enrolling and obtaining credit should be made prior to undertaking an internship. Consent of instructor
Repeat for Credit
Y
Typically Offered
Fall, Spring, Summer
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BIOL 696 - Fisheries Management and Techniques Credits 4
Historical and contemporary issues in the management and conservation of exploited fishes. Methods for managing fisheries resources in streams, lakes, and ponds including estimating abundances, quantifying age and growth, manipulating populations, modeling population dynamics, culturing fishes, and improving aquatic habitat.
Note
Three hours lecture and three hours lab per week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 221 and BIOL 333.
Typically Offered
Fall-Even Years
-
BIOL 697 - Topics in Biology Credits 1-6
Special course offering in an area of faculty expertise and/or supervised independent study project.
Note
Consent of instructor
Repeat for Credit
Y
Typically Offered
Fall, Spring, Summer
-
BIOL 698 - Research in Biology Credits 0-8
Undergraduate research project pursued under the direction of a faculty mentor. A minimum of 45 hours of research effort is expected over the semester for each credit hour the student is enrolled.
Note
Consent of instructor
Repeat for Credit
Y
Typically Offered
Fall, Spring, Summer
-
BIOL 699 - Undergraduate Seminar in Biology Credits 1
Attendance at seminars or journal clubs with topics of a biological nature. A summary and personal reflection on each meeting is submitted for course credit.
Note
Consent of instructor
Repeat for Credit
Y
Typically Offered
Fall, Spring
-
BIOL 705 - Eukaryotic Genetics Credits 3
An integrated exploration of transmission genetics and molecular genetics of eukaryotic organisms. The focus will be on genetic model organisms and their contributions to our understanding of mechanisms of genetic transmission and exchange, mutagenesis, gene expression, and regulation of cell division and development. Modern approaches to genomic analysis will be discussed.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 398 and BIOCH 521.
Typically Offered
Fall
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BIOL 707 - Advanced Cell Biology Credits 3
Selected current topics in cell biology which reflect recent advances in the field. Major topics include membranes and transport, protein sorting, signal transduction, cell adhesion and motility, cell cycle, apoptosis, and specialized cell functions.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 541.
Typically Offered
Fall
-
BIOL 730 - General Virology Credits 3
Theoretical and experimental basis of virology, with emphasis on how viruses manipulate cells to insure their own survival, mechanisms of virus replication, principles of virus host interactions including how viruses cause disease, and selected medically relevant viruses.
Repeat for Credit
N
Requisites
Prerequisite: Twelve hours of biological sciences, including BIOL 398 and BIOL 255; and BIOCH 521 or equivalent; consent of instructor.
Typically Offered
Spring
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BIOL 734 - Introduction to Genomics and Bioinformatics Credits 4
A problem solving approach to understanding genomics and bioinformatics. Practical use of databases and web-based tools used to study biological problems. Introduction to the algorithms behind these tools.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 398 or CIS 575.
Typically Offered
Other
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BIOL 743 - Advanced Ornithology Credits 3
Organized conceptually around the rapid decline of the continent’s avifauna. Following an introductory module introducing the declines, the material is put in context, covering historic and geographic contexts, morphology and physiology, our local avifauna, tools and approaches in ornithology, birds in a changing world, behavioral diversity, and solutions to the problems faced by birds, globally.
Repeat for Credit
N
Requisites
Prerequisite: Open to all students enrolled in MS or PhD program in Biology or related fields.
Typically Offered
Spring
-
BIOL 805 - Advanced Mycology Credits 3
Study of fungi, with emphasis on structure, identification, classification, phylogeny, and economic importance.
Note
One hour lecture and six hours lab a week.
Repeat for Credit
Y
Requisites
Prerequisite: BIOL 604.
Typically Offered
Spring-Odd Years
-
BIOL 807 - Human Genetics Credits 3
An exploration of human genetics from a molecular perspective, grounded in clinical applications. Provides theoretical knowledge and practical skills in human genome analysis, emphasizing the analysis of case studies and molecular diagnostics techniques. Through a combination of lectures, hands-on laboratory exercises, and discussions, students will examine the implications of genetics and genomics for medicine, clinical diagnostics, genetic counseling, and related fields.
Note
One hour of lecture and two two-hour lecture/labs per week.
Repeat for Credit
N
Requisites
Prerequisite: A genetics course and graduate standing.
Typically Offered
Spring
-
BIOL 808 - Mechanisms of Gene Regulation Credits 3
Examines important concepts in the mechanisms of gene regulation at the molecular and atomic levels. Focuses on gene regulation at the level of transcription, splicing and translation and on diverse roles of RNA in catalysis, signaling and innate immunity.
Note
Composed of a lecture (2 hr) covering important concepts and a student recitation (1 hr) of scientific articles.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 541, or BIOL 705, or BIOL 707.
Typically Offered
Spring
-
BIOL 810 - Analysis of Ecological Gradients Credits 3
An introduction to analytical methods and conceptual approaches to evaluate patterns of communities across environmental gradients. Multivariate statistical techniques will be used to analyze data and quantify species abundance patterns in a variety of environments.
Repeat for Credit
Y
Typically Offered
Spring
-
BIOL 818 - Advanced Aquatic Ecology Credits 3
A study of advanced issues adn methodology in limnological sciences, including a workship on algal taxonomy, and an applied group project.
Note
Two lecture hours, two laboratory hours per week.
Repeat for Credit
Y
Typically Offered
Fall-Odd Years
-
BIOL 822 - Landscape Ecology Credits 4
Effect of spatial pattern on ecological processes. Course will emphasize how spatial complexity emerges and is maintained in ecological systems, the analysis of spatial pattern, scaling issues, the ecological consequences of spatial pattern and applications for conservation and ecosystem management in both aquatic and terrestrial systems.
Note
Two lecture hours, one discussion hour and three laboratory hours per week.
Repeat for Credit
Y
Requisites
Prerequisite: BIOL 529.
Typically Offered
Fall-Odd Years
-
BIOL 823 - Demographic Methods Credits 3
Theory and methods of quantitative approaches for the study of population dynamics. Advances in matrix methods and mark-recapture statistics will be emphasized.
Note
Two hours lecture and one hour studio per week.
Repeat for Credit
Y
Requisites
Prerequisite: BIOL 684 or BIOL 696.
Typically Offered
Fall-Even Years
-
BIOL 826 - Nutrient Dynamics Credits 3
The cycling of elements in ecosystems with emphasis on macronutrients such as nitrogen, phosphorous, and major cations, and the influence of variables such as acid rain on nutrient dynamics.
Note
Three hours lecture per week.
Repeat for Credit
N
Requisites
Prerequisite: BIOL 529 and CHM 210.
Typically Offered
Spring-Odd Years
-
BIOL 830 - Advanced Virology Credits 3
In-depth readings and discussions of current topics in virology and the impact of viruses in medicine and biotechnology, with emphasis on recent research literature.
Repeat for Credit
Y
Requisites
Prerequisite: BIOL 730 or consent of instructor.
Typically Offered
Spring-Odd Years
-
BIOL 831 - Host-Pathogen Interactions Credits 3
Explores fundamental aspects of the mechanisms, ecology, and evolution of host-pathogen interactions. Students will have the opportunity to gain in-depth knowledge of these topics through reading, discussion, presentation, and writing exercises.
Repeat for Credit
N
Requisites
Prerequisite: Advanced undergraduate-level understanding of molecular biology/cell biology/ecology/evolution.
Typically Offered
Fall-Even Years
-
BIOL 835 - Cellular and Molecular Parasitology Credits 3
Biochemistry, immunology, and molecular biology of medically important eukaryotic parasites.
Note
Three hours lecture
Repeat for Credit
Y
Requisites
Prerequisite: BIOCH 521 or equivalent.
Typically Offered
On sufficient demand
-
BIOL 840 - Molecular and Cellular Immunology Credits 3
Discussions and readings covering the molecular and cellular interactions during innate and adaptive immune responses.
Repeat for Credit
Y
Requisites
Prerequisite: BIOL 670.
Typically Offered
Fall-Odd Years
-
BIOL 850 - Advanced Topics in Immunology Credits 1-2
Current research in immunology.
Repeat for Credit
Y
Requisites
Prerequisite: BIOL 670 and consent of instructor.
Typically Offered
On sufficient demand
-
BIOL 855 - Molecular Biology of Cellular Membranes Credits 3
Intensive discussions of molecular interactions between proteins and lipids within cellular membranes, and the impact of these interactions on diverse cellular functions. Format is primarily group discussion of primary literature, with the goal of instilling the ability to think critically and evaluate published studies.
Repeat for Credit
Y
Requisites
Prerequisite: BIOL 541 and BIOCH 521.
Typically Offered
Fall
-
BIOL 860 - Modern Molecular Approaches Credits 3
An introduction to approaches and techniques used in the molecular analysis of biological systems, with a focus on modern molecular and cellular biological approaches. Format is primarily group discussion of primary literature, with the goal of instilling the ability to think critically and evaluate published studies.
Repeat for Credit
Y
Requisites
Prerequisite: BIOL 705 and BIOL 707.
Typically Offered
Spring
-
BIOL 863 - Professional Skills and Ethics Credits 3
An introduction for graduate students in the mechanics of becoming a scientist and professional biologist. Students actively participate in learning professional skills such as proposal writing and reviewing, professional oral and poster presentations,communicating science to the general public, abstract and manuscript preparation, and other topics. Ethical conduct in research and professional settings is discussed throughout the course.
Note
Offered every spring, meets twice a week for 1.5 hours. Required course for biology graduate students.
Repeat for Credit
Y
Typically Offered
Spring
-
BIOL 865 - Advanced Plant Ecology Credits 4
Advanced study of theory in population and community ecology as applied to higher plants.
Note
Emphasis on current research in plant population ecology, species interactions, community structure, and four hours lecture/recitation per week.
Repeat for Credit
Y
Requisites
Prerequisite: BIOL 529 or a plant ecology course.
Typically Offered
Fall-Even Years
-
BIOL 868 - Advanced Cellular and Developmental Biology Credits 3
Chemistry, structure, and function of cellular systems in growth, development, and reproduction.
Repeat for Credit
Y
Requisites
Prerequisite: BIOCH 755 or equivalent.
Typically Offered
Fall-Odd Years
-
BIOL 870 - Advanced Plant Systematics Credits 4
Taxonomy, phylogenetic inference and major themes in the evolution of vascular plants.
Note
Two hours lecture, one hour discussion, and three hours lab per week. Lab format will include individual or small group research projects.
Repeat for Credit
Y
Requisites
Prerequisite: BIOL 551 and consent of instructor.
Typically Offered
Spring-Even Years
-
BIOL 875 - Evolutionary Ecology Credits 3
A study of the evolution of population, community, and ecosystem structure.
Note
Two hours lecture and one hour recitation a week.
Repeat for Credit
Y
Requisites
Prerequisite: BIOL 529.
Typically Offered
Spring-Odd Years
-
BIOL 886 - Confocal, Fluorescence and Light Microscopy Credits 3
An introduction to theories, functions and applications of confocal, fluorescence and light microscopy, and fluorescent molecules. Lab emphasis on students working on independent research projects requiring microscopy.
Note
Two hours of lecture and three hours of lab per week.
Repeat for Credit
Y
Typically Offered
Fall-Odd Years
-
BIOL 888 - Electron Microscopy Techniques Credits 3
Theory and techniques involved in using the transmission electron microscope for the study of biological materials. Includes individualized instruction on the operation of the Philips 201 electron microscope and techniques for processing biological samples.
Repeat for Credit
Y
Requisites
Prerequisite: Current participation in research requiring electron microscope.
Typically Offered
Fall-Even Years
-
BIOL 890 - Advanced Topics in Biology Credits 1-6
Advanced biology.
Note
Consent of instructor
Repeat for Credit
Y
Typically Offered
Fall, Spring, Summer
-
BIOL 891 - Advanced Problems in Biology Credits 1-8
Advanced Problems in Biology
Note
Consent of instructor
Repeat for Credit
Y
Typically Offered
Fall, Spring, Summer
-
BIOL 895 - Graduate Seminar in Biology Credits 0-1
Attendance/participation in departmental research-related seminars.
Repeat for Credit
Y
Typically Offered
Fall, Spring
-
BIOL 898 - Master’s Research in Biology Credits 1-9
Master’s Research in Biology
Repeat for Credit
Y
Typically Offered
Fall, Spring, Summer
-
BIOL 997 - Postdoctoral Research in Biology Credits 1-12
Advanced-level research in collaboration with a faculty member, involving projects in any area of biology.
Repeat for Credit
Y
Requisites
Prerequisite: Ph.D. degree or equivalent.
Typically Offered
Fall, Spring, Summer
-
BIOL 998 - PhD Research in Biology Credits 1-18
PhD Research in Biology
Repeat for Credit
Y
Typically Offered
Fall, Spring, Summer
-
BIOL 999 - PhD Research in Microbiology Credits 1-18
PhD Research in Microbiology
Repeat for Credit
Y
Typically Offered
Fall, Spring, Summer
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